William Hope / Edward (US) /  Toby (US) /  The Duke of Boxford (US)(voice)
Steven Kynman / Porter /  Ryan /  Paxton /  Dart (voice)
Joseph May / Thomas (US) (voice)
David Menkin / Porter (US) (voice)
Joe Mills / Douglas (UK & US) (voice)
Tracy-Ann Oberman / Daisy (UK & US) (voice)
Nigel Pilkington / Percy (UK) (voice)
Rob Rackstraw / James (UK & US) /  Toby (UK) /  Donald (UK & US) /  Mr. Fergus Duncan (UK & US) /  The Thin Clergyman (UK & US)(voice)
Christopher Ragland / Percy (US) /  Troublesome Trucks (UK & US) (voice)
Miranda Raison / Millie (UK & US) (voice)
Kerry Shale / Henry (US) /  Gordon (US) /  Diesel (UK & US) /  Harold (US)(voice)
Tom Stourton / Rex (UK & US) /  Terence (UK & US) /  The Fat Clergyman (UK & US) (voice)
Tim Whitnall / Mike (UK & US) /  Jerome (UK & US) (voice)
Keith Wickham / Edward (UK) /  Henry (UK) /  Gordon (UK) /  James (UK) / Salty (UK & US) /  Den (UK & US) /  Stafford (UK & US) / Bert (UK & US) / Harold (UK) /  Bertie (UK & US) (voice)
Keith Wickham / Sir Topham Hatt (UK & US) /  Dowager Hatt (UK & US) /  Farmer Finney (UK & US) /  The Knapford Stationmaster (UK & US)/  The Signalman (UK & US) (voice)
Keith Wickham / The Workmen (UK & US) /  The Woodsman (UK & US) /  The Carollers (UK & US) /  The Grumpy Passenger (UK & US) /  Passengers (UK & US) (voice)
Matt Wilkinson / Spencer (UK) /  Cranky (UK) /  The Duke of Boxford (UK) / Bert's Driver (UK & US) /  Rex's Driver (UK & US) (voice)
Glenn Wrage / Spencer (US) /  Cranky (US) (voice)
Ben Forster / Mr. Perkins

Maggie Ollerenshaw / Henrietta (UK & US) (voice)
Rob Rackstraw / James (US) /  Donald (UK & US) /  The Bird Watcher (UK & US) /  Douglas' Driver (UK & US) /  Passengers (UK & US) / Landmarks of Sodor Announcer (voice)
Christopher Ragland / Percy (US) (voice)
Miranda Raison / Millie (UK & US) (voice)
Kerry Shale / Gordon (US) /  Diesel (UK & US) /  Harold (US) (voice)
Keith Wickham / Glynn (UK & US) /  Sir Topham Hatt (UK & US) (voice)